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The above part, made me think...but Hooper is so scared of other people that he hides the entire time so that won't help.
I remember you mentioning that Hooper was scared of people at first...but I thought it had to do with 'all the things in the house...being new to him." And naturally some cats will run and hide whenever new people come into their territory. Such as that repair person.
But now I am thinking that if Hooper has a stronger 'flight response'...versus ...a 'fight response'...then you could possibly use this to your advantage.
I sometimes think that having one cat dominant...and one submissive....tends to work out better, as opposed to both wanting to be dominant, and choosing to 'fight'. (Though I wouldn't want the submissive one to get bullied on, either.)
In this way, I tend to see Hooper as desperately wanting to play with Twyla, and not actually wanting to hurt her.
Though I do understand, that Twyla is a senior cat, and cannot keep up, or even want to play, at the same level as Hooper.
Not to mention, that Hooper's energy level, would just annoy Twyla.
Except that Twyla does go after Hooper, which makes me think ....that she does want to 'smack, swat, rough him up a little'.
Perhaps when she fell over, it was her going to 'ground' ...and thereby wanting to use her 'back legs' to kick him off. More in a play tussle, than an actual rough fight. It is really difficult to distinguish between the two, since cats move so lightning fast, and end up separating just as fast...if no humans intervene.
It's so hard though, from where we are sitting, to judge that.
Anyhow, my point was only ...that if Hooper still 'runs away'...'flight'...then he may...still respond well to Twyla's cues.
Has Twyla's growling lessened, at this point in time, as opposed to how she was in those first few videos?It was mostly Hooper going after Twyla, but she went after him a little too. It was about ten minutes tops.
If they have, then I would see that as progress.